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We will continue to update this list as new research is published.

Conservation methods
Bertolino, Sandro & Lurz, Peter & Shuttleworth, Craig & Martinoli, Adriano & Wauters, Lucas. (2016). ‘25. The management of grey squirrel populations in Europe: evolving best practice The management of invasive species’ in Shuttleworth, C., Lurz, P., Gurnell, J. (eds.) The Grey Squirrel Ecology & Management of an Invasive Species in Europe. European Squirrel Initiative pp.495-516.

Dunn M., Marzano M., Forster J (2018) Public attitudes towards “pest” management: Perceptions on squirrel management strategies in the UK. Biological Conservation 222:52-63. https://doi.org/10.1016/j.biocon.2018.03.020 

IUCN/SSC (2013) Guidelines for Reintroductions and Other Conservation
Version 1.0. Gland, Switzerland: IUCN Species Survival Commission,
viiii + 57 pp. Available at: https://portals.iucn.org/library/sites/library/files/documents/2013-009.pdf

Jones, H., White, A., Lurz, P. and Shuttleworth, C., (2017) Mathematical models for invasive species management: Grey squirrel control on Anglesey. Ecological Modelling, 359, pp.276-284. https://doi.org/10.1016/j.ecolmodel.2017.05.020 

Ogden R., Shuttleworth C., McEwing R., et al., (2005) Genetic management of the red squirrel, Sciurus vulgaris: a practical approach to regional conservation. Conservation Genetics, 6(4):511-525. https://doi.org/10.1007/s10592-005-9006-8

Red Squirrel Survival Trust (2021) Saving the Red Squirrel: Landscape Scale Recovery. Edited by Shuttleworth, C.M., Lurz, P.W., & Robinson, N. Available at: https://www.rsst.org.uk/wp-content/uploads/2021/03/SavingRS2021Final.pdf

Schuchert, P., Shuttleworth, C.M., McInnes, C.J., Everest, D.J., & Rushton, S.P., (2012) Shared Experience of Red Squirrel Conservation Practice. Edited by Shuttleworth, C.M., Lurz, P.W., Halliwell, E.C. Available at: https://www.europeansquirrelinitiative.org/wp-content/uploads/2016/11/Red-Squirrel-Conservation-eBook.pdf

Schuchert, P., Shuttleworth, C. M., McInnes, C. J., Everest, D. J., & Rushton, S. P. (2014). Landscape scale impacts of culling upon a European grey squirrel population: can trapping reduce population size and decrease the threat of squirrelpox virus infection for the native red squirrel?. Biological invasions16(11), 2381-2391. https://doi.org/10.1007/s10530-014-0671-8

Shuttleworth, C.M., Robinson, N., Halliwell, E.C., Clews-Roberts, R., Peek, H., Podgornik, G., Stinson, M., Rice, S., Finlay, C., McKinney, C. and Everest, D.J., (2020) Evolving grey squirrel management techniques in Europe. Management of Biological Invasions, 11(4), p.747. https://doi.org/10.3391/mbi.2020.11.4.09 

Signorile, A.L. and Shuttleworth, C.M., (2016) Genetic evidence of the effectiveness of grey squirrel control operations: lessons from the Isle of Anglesey. The grey squirrel—ecology & management of an invasive species in Europe. European Squirrel Initiative, Woodbridge, pp.439-451.

White, A. and Lurz, P.W.W., (2018) Grey squirrel control in southern Scotland: A model analysis. Forest Enterprise Scotland.

Habitat & Diet
Babińska-Werka, J. and Żółw, M., (2008) Urban populations of the red squirrel (Sciurus vulgaris) in Warsaw. In Annales Zoologici Fennici (Vol. 45, No. 4, pp. 270-276). Finnish Zoological and Botanical Publishing Board. https://doi.org/10.5735/086.045.0405  

Haigh A., Butler F., O’Riordan R., et al., (2017) Managed parks as a refuge for the threatened red squirrel (Sciurus vulgaris) in light of human disturbance. Biological Conservation, 211:29-36. https://doi.org/10.1016/j.biocon.2017.05.008 

Jokimäki J., Selonen V., Lehikoinen A., et al., (2017) The role of urban habitats in the abundance of red squirrels (Sciurus vulgaris, L.) in Finland. Urban Forestry & Urban Greening, 27:100-108. https://doi.org/10.1016/j.ufug.2017.06.021 

Krauze-Gryz, D. and Gryz, J., (2015). ‘A review of the diet of the red squirrel (Sciurus vulgaris) in different types of habitats’ in CM Shuttleworth, PWW Lurz, MW Hayward (eds.) Red squirrels: ecology, conservation & management in Europe. European Squirrel Initiative, (s 3950).
Rézouki C., Dozières A., Le Cœur C., et al., (2014) A viable population of the European red squirrel in an urban park. PloS one, (8):105-111. https://doi.org/10.1371/journal.pone.0105111 

Virus & disease
Bruemmer, C.M., Rushton, S.P., Gurnell, J., Lurz, P.W.W., Nettleton, P., Sainsbury, A.W., Duff, J.P., Gilray, J. and McInnes, C.J., (2010) Epidemiology of squirrelpox virus in grey squirrels in the UK. Epidemiology & Infection, 138(7), pp.941-950. https://doi.org/10.1017/S0950268810000816 

Chantrey, J., Dale, T.D., Read, J.M., White, S., Whitfield, F., Jones, D., McInnes, C.J. and Begon, M., (2014) European red squirrel population dynamics driven by squirrelpox at a gray squirrel invasion interface. Ecology and Evolution, 4(19), pp.3788-3799. https://doi.org/10.1002/ece3.1216 

Chantrey, J., Dale, T., Jones, D., Begon, M. and Fenton, A., 2019. The drivers of squirrelpox virus dynamics in its grey squirrel reservoir host. Epidemics, 28, p.100352. https://doi.org/10.1016/j.epidem.2019.100352 

Collins, L.M., Warnock, N.D., Tosh, D.G., McInnes, C., Everest, D., Montgomery, W.I., Scantlebury, M., Marks, N., Dick, J.T. and Reid, N., (2014) Squirrelpox virus: assessing prevalence, transmission and environmental degradation. PLoS One, 9(2), p.e89521. https://doi.org/10.1371/journal.pone.0089521 

Everest D., Shuttleworth C., Stidworthy MF., et al., (2014) Adenovirus: an emerging factor in red squirrel Sciurus vulgaris conservation. Mammal Review, 44(3-4):225-233. https://doi.org/10.1111/mam.12025 

Romeo, C., Ferrari, N., Lanfranchi, P., Saino, N., Santicchia, F., Martinoli, A. and Wauters, L.A., (2015) Biodiversity threats from outside to inside: effects of alien grey squirrel (Sciurus carolinensis) on helminth community of native red squirrel (Sciurus vulgaris). Parasitology research, 114(7), pp.2621-2628. https://doi.org/10.1007/s00436-015-4466-3 

Romeo C., Pisanu B., Ferrari N., et al., (2013) Macroparasite community of the Eurasian red squirrel (Sciurus vulgaris): poor species richness and diversity. Parasitol Res, 112(10):3527-353. https://doi.org/10.1007/s00436-013-3535-8 

Rushton, S.P., Lurz, P.W.W., Gurnell, J., Nettleton, P., Bruemmer, C., Shirley, M.D.F. and Sainsbury, A.W., (2006) Disease threats posed by alien species: the role of a poxvirus in the decline of the native red squirrel in Britain. Epidemiology & Infection, 134(3), pp.521-533. https://doi.org/10.1017/S0950268805005303 

Shuttleworth, C.M., Everest, D., Holmes, P., Bell, S., Cripps, R. (2022) An Opportunistic Assessment of the Impact of Squirrelpox Disease Outbreaks upon a Red Squirrel Population Sympatric with Grey Squirrels in Wales. Animals12, 99. https://doi.org/10.3390/ani12010099

White, A., Lurz, P.W., Bryce, J., Tonkin, M., Ramoo, K., Bamforth, L., Jarrott, A. and Boots, M., (2016) Modelling disease spread in real landscapes: Squirrelpox spread in Southern Scotland as a case study. https://doi.org/10.4404/hystrix-27.1-11657 

Red & grey squirrels
Chow P., Lurz P., Lea S. (2018) A battle of wits? Problem-solving abilities in invasive eastern grey squirrels and native Eurasian red squirrels. Animal Behaviour, (137):11-20. https://doi.org/10.1016/j.anbehav.2017.12.022 

Gurnell, J., Wauters, L.A., Lurz, P.W. and Tosi, G., (2004) Alien species and interspecific competition: effects of introduced eastern grey squirrels on red squirrel population dynamics. Journal of Animal Ecology, 73(1), pp.26-35.  https://doi.org/10.1111/j.1365-2656.2004.00791.x 

Lurz, P.W.W., Rushton, S.P., Wauters, L.A., Bertolino, S., Currado, I., Mazzoglio, P. and Shirley, M.D.F., (2001) Predicting grey squirrel expansion in North Italy: a spatially explicit modelling approach. Landscape Ecology, 16(5), pp.407-420. https://doi.org/10.1023/A:1017508711713 

Sandro, B., (2008) Introduction of the American grey squirrel (Sciurus carolinensis) in Europe: a case study in biological invasion. Current Science, pp.903-906. www.jstor.org/stable/24103188.