We will continue to update this list as new research is published.
Conservation methods |
Bertolino, Sandro & Lurz, Peter & Shuttleworth, Craig & Martinoli, Adriano & Wauters, Lucas. (2016). ‘25. The management of grey squirrel populations in Europe: evolving best practice The management of invasive species’ in Shuttleworth, C., Lurz, P., Gurnell, J. (eds.) The Grey Squirrel Ecology & Management of an Invasive Species in Europe. European Squirrel Initiative pp.495-516. |
Dunn M., Marzano M., Forster J (2018) Public attitudes towards “pest” management: Perceptions on squirrel management strategies in the UK. Biological Conservation 222:52-63. https://doi.org/10.1016/j.biocon.2018.03.020 |
IUCN/SSC (2013) Guidelines for Reintroductions and Other Conservation Translocations. Version 1.0. Gland, Switzerland: IUCN Species Survival Commission, viiii + 57 pp. Available at: https://portals.iucn.org/library/sites/library/files/documents/2013-009.pdf |
Jones, H., White, A., Lurz, P. and Shuttleworth, C., (2017) Mathematical models for invasive species management: Grey squirrel control on Anglesey. Ecological Modelling, 359, pp.276-284. https://doi.org/10.1016/j.ecolmodel.2017.05.020 |
Ogden R., Shuttleworth C., McEwing R., et al., (2005) Genetic management of the red squirrel, Sciurus vulgaris: a practical approach to regional conservation. Conservation Genetics, 6(4):511-525. https://doi.org/10.1007/s10592-005-9006-8 |
Red Squirrel Survival Trust (2021) Saving the Red Squirrel: Landscape Scale Recovery. Edited by Shuttleworth, C.M., Lurz, P.W., & Robinson, N. Available at: https://www.rsst.org.uk/wp-content/uploads/2021/03/SavingRS2021Final.pdf |
Schuchert, P., Shuttleworth, C.M., McInnes, C.J., Everest, D.J., & Rushton, S.P., (2012) Shared Experience of Red Squirrel Conservation Practice. Edited by Shuttleworth, C.M., Lurz, P.W., Halliwell, E.C. Available at: https://www.europeansquirrelinitiative.org/wp-content/uploads/2016/11/Red-Squirrel-Conservation-eBook.pdf |
Schuchert, P., Shuttleworth, C. M., McInnes, C. J., Everest, D. J., & Rushton, S. P. (2014). Landscape scale impacts of culling upon a European grey squirrel population: can trapping reduce population size and decrease the threat of squirrelpox virus infection for the native red squirrel?. Biological invasions, 16(11), 2381-2391. https://doi.org/10.1007/s10530-014-0671-8 |
Shuttleworth, C.M., Robinson, N., Halliwell, E.C., Clews-Roberts, R., Peek, H., Podgornik, G., Stinson, M., Rice, S., Finlay, C., McKinney, C. and Everest, D.J., (2020) Evolving grey squirrel management techniques in Europe. Management of Biological Invasions, 11(4), p.747. https://doi.org/10.3391/mbi.2020.11.4.09 |
Signorile, A.L. and Shuttleworth, C.M., (2016) Genetic evidence of the effectiveness of grey squirrel control operations: lessons from the Isle of Anglesey. The grey squirrel—ecology & management of an invasive species in Europe. European Squirrel Initiative, Woodbridge, pp.439-451. |
White, A. and Lurz, P.W.W., (2018) Grey squirrel control in southern Scotland: A model analysis. Forest Enterprise Scotland. |
Habitat & Diet |
Babińska-Werka, J. and Żółw, M., (2008) Urban populations of the red squirrel (Sciurus vulgaris) in Warsaw. In Annales Zoologici Fennici (Vol. 45, No. 4, pp. 270-276). Finnish Zoological and Botanical Publishing Board. https://doi.org/10.5735/086.045.0405 |
Haigh A., Butler F., O’Riordan R., et al., (2017) Managed parks as a refuge for the threatened red squirrel (Sciurus vulgaris) in light of human disturbance. Biological Conservation, 211:29-36. https://doi.org/10.1016/j.biocon.2017.05.008 |
Jokimäki J., Selonen V., Lehikoinen A., et al., (2017) The role of urban habitats in the abundance of red squirrels (Sciurus vulgaris, L.) in Finland. Urban Forestry & Urban Greening, 27:100-108. https://doi.org/10.1016/j.ufug.2017.06.021 |
Krauze-Gryz, D. and Gryz, J., (2015). ‘A review of the diet of the red squirrel (Sciurus vulgaris) in different types of habitats’ in CM Shuttleworth, PWW Lurz, MW Hayward (eds.) Red squirrels: ecology, conservation & management in Europe. European Squirrel Initiative, (s 3950). |
Rézouki C., Dozières A., Le Cœur C., et al., (2014) A viable population of the European red squirrel in an urban park. PloS one, (8):105-111. https://doi.org/10.1371/journal.pone.0105111 |
Virus & disease |
Bruemmer, C.M., Rushton, S.P., Gurnell, J., Lurz, P.W.W., Nettleton, P., Sainsbury, A.W., Duff, J.P., Gilray, J. and McInnes, C.J., (2010) Epidemiology of squirrelpox virus in grey squirrels in the UK. Epidemiology & Infection, 138(7), pp.941-950. https://doi.org/10.1017/S0950268810000816 |
Chantrey, J., Dale, T.D., Read, J.M., White, S., Whitfield, F., Jones, D., McInnes, C.J. and Begon, M., (2014) European red squirrel population dynamics driven by squirrelpox at a gray squirrel invasion interface. Ecology and Evolution, 4(19), pp.3788-3799. https://doi.org/10.1002/ece3.1216 |
Chantrey, J., Dale, T., Jones, D., Begon, M. and Fenton, A., 2019. The drivers of squirrelpox virus dynamics in its grey squirrel reservoir host. Epidemics, 28, p.100352. https://doi.org/10.1016/j.epidem.2019.100352 |
Collins, L.M., Warnock, N.D., Tosh, D.G., McInnes, C., Everest, D., Montgomery, W.I., Scantlebury, M., Marks, N., Dick, J.T. and Reid, N., (2014) Squirrelpox virus: assessing prevalence, transmission and environmental degradation. PLoS One, 9(2), p.e89521. https://doi.org/10.1371/journal.pone.0089521 |
Everest D., Shuttleworth C., Stidworthy MF., et al., (2014) Adenovirus: an emerging factor in red squirrel Sciurus vulgaris conservation. Mammal Review, 44(3-4):225-233. https://doi.org/10.1111/mam.12025 |
Romeo, C., Ferrari, N., Lanfranchi, P., Saino, N., Santicchia, F., Martinoli, A. and Wauters, L.A., (2015) Biodiversity threats from outside to inside: effects of alien grey squirrel (Sciurus carolinensis) on helminth community of native red squirrel (Sciurus vulgaris). Parasitology research, 114(7), pp.2621-2628. https://doi.org/10.1007/s00436-015-4466-3 |
Romeo C., Pisanu B., Ferrari N., et al., (2013) Macroparasite community of the Eurasian red squirrel (Sciurus vulgaris): poor species richness and diversity. Parasitol Res, 112(10):3527-353. https://doi.org/10.1007/s00436-013-3535-8 |
Rushton, S.P., Lurz, P.W.W., Gurnell, J., Nettleton, P., Bruemmer, C., Shirley, M.D.F. and Sainsbury, A.W., (2006) Disease threats posed by alien species: the role of a poxvirus in the decline of the native red squirrel in Britain. Epidemiology & Infection, 134(3), pp.521-533. https://doi.org/10.1017/S0950268805005303 |
Shuttleworth, C.M., Everest, D., Holmes, P., Bell, S., Cripps, R. (2022) An Opportunistic Assessment of the Impact of Squirrelpox Disease Outbreaks upon a Red Squirrel Population Sympatric with Grey Squirrels in Wales. Animals, 12, 99. https://doi.org/10.3390/ani12010099 |
White, A., Lurz, P.W., Bryce, J., Tonkin, M., Ramoo, K., Bamforth, L., Jarrott, A. and Boots, M., (2016) Modelling disease spread in real landscapes: Squirrelpox spread in Southern Scotland as a case study. https://doi.org/10.4404/hystrix-27.1-11657 |
Red & grey squirrels |
Chow P., Lurz P., Lea S. (2018) A battle of wits? Problem-solving abilities in invasive eastern grey squirrels and native Eurasian red squirrels. Animal Behaviour, (137):11-20. https://doi.org/10.1016/j.anbehav.2017.12.022 |
Gurnell, J., Wauters, L.A., Lurz, P.W. and Tosi, G., (2004) Alien species and interspecific competition: effects of introduced eastern grey squirrels on red squirrel population dynamics. Journal of Animal Ecology, 73(1), pp.26-35. https://doi.org/10.1111/j.1365-2656.2004.00791.x |
Lurz, P.W.W., Rushton, S.P., Wauters, L.A., Bertolino, S., Currado, I., Mazzoglio, P. and Shirley, M.D.F., (2001) Predicting grey squirrel expansion in North Italy: a spatially explicit modelling approach. Landscape Ecology, 16(5), pp.407-420. https://doi.org/10.1023/A:1017508711713 |
Sandro, B., (2008) Introduction of the American grey squirrel (Sciurus carolinensis) in Europe: a case study in biological invasion. Current Science, pp.903-906. www.jstor.org/stable/24103188. |